[S]ymptoms included interveinal darkening, some puckering, crinkling and irregularly thickened leaves. Mature forest trees exhibited thinned crowns and branch dieback. ... Nearly angular leaf spots, upon close inspection with a stereo-microscope, showed very small angular lesions within the darkened areas ... BLD is associated with tree mortality within 7 years of detection.

- Carta, Lynn Kay; Handoo, Zafar A.; Li, Shiguang; Kantor, Mihail; Bauchan, Gary; McCann, David; Gabriel, Colette K.; Yu, Qing; Reed, Sharon; Koch, Jennifer; Martin, Danielle; Burke, David J.: (2020) Beech leaf disease symptoms caused by newly recognized nematode subspecies Litylenchus crenatae mccannii (Anguinata) described from Fagus grandifolia in North America.^©